Herpetology Notes

About the Journal

Herpetology Notes is an online-only and Open Access journal (under CC BY-NC 4.0) published by the Societas Europaea Herpetologica (SEH) alongside its flagship journal, Amphibia-Reptilia. Processing, editing, and typesetting of manuscripts is carried out by an international editorial team. The current rejection rate is around 30%.

Announcements

Welcome to the NEW Herpetology Notes website!

2024-12-01

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Current Issue

Vol. 18 (2025)

Published: 2025-03-05

Articles

First record of Trimerodytes praemaxillaris (Angel, 1929) from Xaisomboun Province, Laos, with additional comments on morphology, natural history, and the type locality

Yanpeng Shen, Christine M. Kaiser, Kanto Nishikawa, Daosavanh Sanamxay, Phouth Inthavong, Hinrich Kaiser

143–153
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Rescue records of Leith’s Sandsnake, Psammophis leithii Günther, 1869, from Surat, Gujarat, India with some additional insights on pholidosis and natural history

Mehul Thakur, Dikansh S. Parmar, Hinrich Kaiser

169–173
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Communal nesting behaviour of female American Alligators, Alligator mississippiensis (Daudin, 1801) in southeastern Oklahoma, USA

Eric C.K. Gren, Jake A. Pruett, Jared P. Wood, Tim M. Patton

215–219
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Tail movements as an anti-predatory behaviour in Hynobius okiensis Sato, 1940

Amaël Borzée, Yucheol Shin, Taiji Shiraishi

225-227
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Six predation events by the Red-banded Snake, Lycodon rufozonatus Cantor, 1842, on Tsushima Island, Japan

Ango Morikawa, Takato Inoue

273-276
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Shedding excess: small tape patches as a lower-impact transmitter attachment method for snakes

Tyler C. Christensen, Jason A. Fantuzzi

307-311
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Bright marks, dark fate: records of predation on Coleodactylus meridionalis Boulenger, 1888 from a movement ecology study in a forest area in southern Bahia, Brazil

Lucca Izaguirres Souza, Charles V. de Jesus Braga, Rafaella Roseno, Luisa M. Diele Viegas, Mirco Solé

351-353
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First record of Kellogg’s Coralsnake, Sinomicrurus kelloggi (Pope, 1928), from Nghe An Province, Vietnam

Shinya Okabe, Dung Van Tran, Kanto Nishikawa, Tao Thien Nguyen, Sinh Van Nguyen, Manh Van Nguyen, Hai Tuan Bui

355-359
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Preliminary data on saltwater crocodiles, Crocodylus porosus Schneider, 1801, and an updated assessment of threats in the Nilwala River, Matara, Sri Lanka

Trevor L. Proctor, Pradeep Rathnasiri, Shaya Honarvar

361-363
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New country records of the Laotian Wolfsnake, Lycodon laoensis Günther, 1864, in Myanmar, discovered via citizen science

Soe Thandar Aung, Pongthep Suwanwaree, Min Thant Aung, Justin M. Bernstein

373–375
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Potential predation by a carabid beetle (Catadromus sp.) larva on an adult Spotted Marsh Frog, Limnodynastes tasmaniensis Günther, 1858, in western Victoria, Australia

Dylan M. Westaway, David A. De Angelis

377-379
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Insights from short-term post-fire monitoring of a population of Hermann’s Tortoise, Testudo hermanni Gmelin, 1789, in Calabria, southern Italy

Francesco L. Leonetti, Antonio G. Adamo, Gianni Giglio, Emilio Sperone

381-385
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Low genetic diversity and relatively strong population genetic structure of Przewalski's Wondergecko, Teratoscincus przewalskii Strauch, 1887, in the Mongolian Gobi Desert

Onolragchaa Ganbold, Erdenetushig Purevee, Tuvshinlkhagva Amartuvshin, Khongorzul Tsagaan, Munguntulga Erdenechuluun, Oyunchuluun Yadamsuren, Suvdtsetseg Chuluunbat, Joon-Woo Lee, Ariunbold Jargalsaikhan

651–658
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Interaction between two Japanese Ratsnakes, Elaphe climacophora (Boie, 1826), at the nest of a Eurasian Wren, Troglodytes troglodytes

Ayaka Soda

463–465
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Decoding the overlap: diagnostic acoustics to distinguish the endangered Magnificent Broodfrog, Pseudophryne covacevichae Ingram & Corben, 1994, from the common Montane Toadlet, Uperoleia altissima Davies et al., 1993 in northern Queensland, Australia

Emily R. Rush

501–511
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Capture events of Forster's Tree Iguana, Liolaemus forsteri Laurent, 1982, by Aplomado Falcon, Falco femoralis Temminck, 1822, in the high Andes of Bolivia

Alejandro Bruno Miranda Calle, Diego Del Carpio Martínez

949-951
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Preliminary assessment of movements and habitat use of the Tabasco Mud Turtle, Kinosternon acutum Gray, 1831, in a tropical rainforest in Belize

Collin McAvinchey , Eric Munscher, Brian Butterfield , Tabitha Hootman, Barney Hall , Rebecca A. Cozad, Thomas Pop, Joseph Pignatelli , Heather Barrett, Jacob Marlin, Zachary Siders , Andrew Walde

891-901
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Nueva información sobre la distribución y biología de la Culebra de Cola Corta Jaspeada, Tantillita brevissima (Taylor, 1937) en México y Guatemala

Emmanuel Javier-Vázquez, Antonio E. Valdenegro-Brito, Juan C. Sánchez-García, Adam G. Clause, Roberto Luna-Reyes, Uri O. García-Vázquez

697–705
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Attempted scavenging and aquatic hunting in the Iberian Asp Viper, Vipera aspis zinnikeri Kramer, 1958

Thierry Gagliano, Matthieu Berroneau, Przemysław Zdunek

989-993
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Life history characteristics of the Colombian Wood Turtle, Rhinoclemmys melanosterna (Gray, 1861), in the middle Magdalena River, Colombia

Vivian P. Páez, Brian C. Bock, Viviana M. Cartagena-Otálvaro

921–930
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The use of fluorescent powdered pigments as a tracking technique for hatchling turtles in Belize

Jessica Henry, Janelle Deneau, Parker Gibbons, Michael Skibsted, Barney Hall, Tom Pop, Zachary A. Siders, Andrew D. Walde, Eric Munscher

455–462
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Male-male combat in free-ranging Falcón Indigo Snakes, Drymarchon caudomaculatus Wüster et al., 2001, in Guajira, Colombia

Lucas S. Almeida, Juliano M.H. Pires, Rodrigo C. Gonzalez

821–824
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Field body temperatures and microclimatic influences in Hermann’s Tortoise, Testudo hermanni Gmelin, 1789, from Thrace

Ceren Nur Özgül, Didem Kurtul, Begüm Boran, Bengi Baycan, Çiğdem Gül, Murat Tosunoğlu

721-730
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First report of consumption of the Italian Three-toed Skink, Chalcides chalcides (Linnaeus, 1758), by a European Pine Marten in Lazio, Italy

Alessia Mariacher, Lorena Di Benedetto, Francesca Maccagnan, Francesco Mariottini, Luca Colla, Sara Tonon, Matteo R. Di Nicola

663–665
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The herpetofauna of the Echinades Island Complex, Ionian Islands, Greece, with notes on the ecological peculiarities of its lizard species

Elias Tzoras, Dionysios Mamasis

759-767
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New records of the Vanzolini’s Spiny-chest Frog, Alsodes vanzolinii (Donoso-Barros, 1974), from south-central Chile

Pedro Pablo Alvarez, Francisco Cifuentes, Álvaro García, Sebastián Barra-Parra, Oscar Hidalgo, Fabián Campos-Cifuentes

787-791
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First photographic evidence of oceanic swimming behaviour in littoral Snake-eyed Skinks, genus Cryptoblepharus, in the Western Indian Ocean

Tim L. Heller, Sohan Sauroy-Toucouère, Kathleen C. Webster, Nassourdine A. Mroudjaé, Hindatou Saidou, Oliver Hawlitschek

659–661
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A noteworthy observation of Corallus caninus (Linnaeus, 1758) within the Corallus batesii (Gray, 1860) range in the Central Brazilian Amazonia

Angel V. Dyugmedzhiev, Kostadin I. Andonov, Vladimir R. Todorov, Nikola M. Stanchev

715-719
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Predation of a White-banded Treefrog, Boana albomarginata (Spix, 1824), by a Parrot Snake, Leptophis liocercus (Wied, 1824), in an Atlantic Forest fragment in Bahia, northeastern Brazil

Tiago Augusto Fonseca Silva, Rebeca Sales dos Santos, Natália de Souza Campos, Arielson dos Santos Protázio

673-675
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Contribution to the knowledge of the gecko Tenuidactylus longipes (Nikolsky, 1896) in northeastern Iran

Saeed S. Hosseinian Yousefkhani, Reza Babaei Savasari

621–624
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Societatis Europeae Herpetologicae brevis historia

Wolfgang Böhme

805–812
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Predation of the Madagascar Dwarf Leaf-toed Gecko, Paroedura vazimba Nussbaum & Raxworthy, 2000, by the Madagascar Bullfrog, Laliostoma labrosum (Cope, 1868), in Ankarafantsika, northwestern Madagascar

Ayane Inoue

605-606
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The wonder of rain: records of Plateau Tiger Salamander, Ambystoma velasci Dugès, 1888, in Salinas, San Luis Potosí, Mexico

Jesús L. Lara-Galván, Jesús Lara-Rayos, Xabier Herrero-Otero, Juan F. Martínez-Montoya, José J. Sigala-Rodríguez

573–576
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First report of piebaldism for the Crowned False Boa, Pseudoboa coronata Schneider, 1801, from the lowlands of Bolivia

Luis R. Rivas, Miguel A. Camacho, Cord B. Eversole, Randy L. Powell

731-733
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Head hiding behaviour in the Eastern Montpellier Snake, Malpolon insignitus fuscus (Fleischmann, 1831)

Alessandro Paterna

607-611
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Tail trifurcation in Hemidactylus aff. malcolmsmithi in southern Mississippi, USA

Jake G. Power, Drew R. Davis

677–678
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Third case of tail bifurcation in Cope’s Mabuya, Notomabuya frenata (Cope, 1863), in Mato Grosso do Sul, Brazil

Nelson R. de Albuquerque, Luciana M. Valério, Roullien H. Martins

467–470
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On the occurrence of a golden colour variant in the Common Spadefoot Toad, Pelobates fuscus (Laurenti, 1768), in the state of Saxony, Germany

Mischa Lauterbach, Judith Adam, Leonard Bolte, Sandra Paule, Timm Reinhardt

941–946
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Limb and digit malformations in the Alpine Newt, Mesotriton alpestris (Laurenti, 1768), in the United Kingdom

Rachel Hester

755-758
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Amplexus between frogs in different families: a microhylid Nilphamarai Narrow-mouthed Frog, Microhyla nilphamariensis Howlader et al., 2015, grasps a Terai Cricket Frog, Minervarya teraiensis (Dubois, 1984), in Nepal

Tapil Prakash Rai, Christine M. Kaiser, Hinrich Kaiser

521–523
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Domestic water buffaloes: an unexpected basking support for two sympatric freshwater turtles, Emys orbicularis (Linnaeus, 1758) and Mauremys rivulata (Valenciennes, 1833) in Türkiye

Ömral Ünsal Özkoç, Cem Akın, Anthony Olivier, Dilara Arslan

839–843
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How the tables may turn: opportunistic scavenging by tadpoles of Discoglossus galganoi Capula et al., 1985 and Epidalea calamita (Laurenti, 1768) on the carcass of a Viperine Watersnake, Natrix maura (Linnaeus, 1758) in northern Portugal

Marisa Naia, Angelica Crottini

813–815
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Unusual use of water-filled tires by Nyctixalus spinosus (Taylor, 1920) in an agroforestry farm in eastern Mindanao, Philippines

Frankie B. M. Juanitas, Dolens J. B. Iñigo, Lief E. D. Gamalo, Aaron F. M. Raganas, Mae A. Responte, Fritzie A. Camino, Charles J. C. Gunay, Marion J. M. M. Achondo

973-976
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Minimum hydroperiod for metamorphosis in the California Tiger Salamander, Ambystoma californiense Gray, 1853

Sean M. O'Brien, Brent P. Helm

613-616
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First record of predation on reptiles by the Sakishima Rice Frog, Fejervarya sakishimensis Matsui et al., 2007, in Japan

Kazuho Anan, Dai Togane, Kohsuke Akita

889-890
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Hidden breeding behaviour in a still undescribed giant Mantidactylus frog from Betampona Strict Nature Reserve, eastern Madagascar

Jean Noel, Karen Freeman, Jean Jacques Jaozandry, Angelica Crottini, Honoré Lava, Georges Rendrirendy, Gonçalo Miranda Rosa, Jean H. Velo, Franco Andreone

969-972
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Post-hatching maternal attendance in wild Burmese Pythons in southern Florida

Jacquelyn C. Guzy, Mark R. Sandfoss, John-Kaarli Rentof, Judith E. Baird-Lujano, Genesis Aponte Santiago, Jose A. Torres, Lisa M. McBride, Matthew F. Metcalf, Amanda M. Kissel, Christina M. Romagosa, Amy A. Yackel Adams, Brandon L. Welty, Melissa A. Miller, Matthew F. McCollister, Frank J. Mazzotti, Kristen M. Hart

849-856
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New records of the Ocellated Skink, Chalcides ocellatus (Forskål, 1775), reveal possible human-mediated translocation to Syros Island, Greece

Taxiarchis Danelis, Petros Stefanou, Apostolos Christopoulos

539-542
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Unusual defensive behaviour combination recorded in Bufotes viridis (Laurenti, 1768)

Daniel Jablonski, Soran H. Ahmed

793-795
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Initial survey of the amphibian chytrid fungus Batrachochytrium dendrobatidis around Bouamir Research Station, Dja Faunal Reserve, Cameroon

Abraham G. Bamba-Kaya, Oscar R. Fokou, Veronica Saenz, Lauren A. Scheinberg, Allison Q. Byrne, LeGrand Nono Gonwouo, C. Guilherme Becker, Rayna C. Bell

529–537
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Updating the snake menu: two cases of ophiophagy by the Western Indigo Snake, Drymarchon corais (Boie, 1827), in a Neotropical rain forest

John Bosu Mensah, Joey Felsch, Ádám Simon, Eva Ringler

981-983
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Ocular anomaly in the Plateau Toad, Anaxyrus compactilis (Wiegmann, 1833), from Jalisco State, West-Central Mexico

Rodolfo Guzmán-Ramírez, Alan I. Olvera-Mendoza, Lizzeth A. Torres-Hernández, César A. Díaz-Marín, Israel Moreno-Lara, Emiliano Hernández-Jiménez, Alexis M. Leonardo-González, Aurelio Ramírez-Bautista

601-603
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New data on introduced geckos of the Mediodactylus kotschyi complex in Hungary

Gergely Babocsay

707-713
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Predation of a Barker Frog, Physalaemus cuvieri Fitzinger, 1826, by a Giant Water Bug, genus Belostoma, in a Cerrado pond Minas Gerais, Brazil

Wendell Sousa-Felix, Rayça Azevedo, Camila R.O. Leal, Priscila S. Carvalho, Diego J. Santana, Karoline Ceron

867–869
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A new record of the Endangered Zolio's Shield-backed Ground Lizard, Philochortus zolii Scortecci, 1934, from Algeria, with genetic insights and a geographic distribution review

Idriss Bouam, D. James Harris, Larbi Tahar-Chaouch

567–571
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Give me five: a case of polymelia in the Common Midwife Toad, Alytes obstetricans (Laurenti, 1768), in Central Spain

Luis Javier Cuéllar, Carlos Caballero-Díaz, Helena Martínez-Gil

617-620
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First record of Paratelmatobius segallai Santos et al., 2019 in the state of São Paulo, southeastern Brazil, based on molecular and morphological evidence

Sandro P. de Faria, Diego J. Santana, Thiago T. Tognolo, Sarah Mângia

861-866
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Don't bother me, I'm venomous: first report of envenoming by an Aesculapian False Coralsnake, Erythrolamprus aesculapii (Linnaeus, 1758), in the Bolivian Yungas

Oliver Quinteros-Muñoz, Edson Pérez

525–527
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New prey record for Baird's Patchnose Snake, Salvadora bairdi Jan, 1860

Irving Y. Rojas-Velasco, Darian A. Neria-Hernández, Luis F. Hidalgo-Licona, O. Iván Martínez Vaca-León

875–877
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New insights into the distribution of Scincus and Scincopus (Scincidae) in Northwest Africa, with the first record of the Sandfish Skink, Scincus scincus (Linnaeus, 1758), from Morocco

Abdellah Bouazza, Gabriel Martínez del Mármol

903-911
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New records of the Golden Spectacled Tegu, Gymnophthalmus speciosus (Hallowell, 1861), in Veracruz, Mexico

Orlando R. Vivanco-Montané, E. Ahmed Bello-Sánchez, Jorge E. Morales-Mávil, Jazmín Enríquez-Roa

883–887
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Prolonged copulation and possible mating plug formation in the Ryukyu Greensnake, Ptyas semicarinata (Hallowell, 1861), on Okinawa Island

Hiroaki Yanagi, Akira Mori

577–579
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Efficacy of GPS-based satellite transmitters to monitor movement and nesting of Alligator Snapping Turtles, Macrochelys temminckii (Troost, 1835)

Christopher Schalk, Jessica Glasscock, Tristan Brownjohn, Jonathan Edwards, Connor Adams

953-961
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Molecular identification of the ant Pheidole nodus in the diet of Indotyphlops braminus (Daudin, 1803) in Taiwan

Chun-Kai Yang, Pei-Lun Sun, Man-Lin Lin, Yi-Ju Yang

871–873
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Use of a handheld metal detector to evaluate metal ingestion in the Alligator Snapping Turtle, Macrochelys temminckii (Troost, 1835)

Kelly L. Garcia, Mandi Gordon, Eric C. Munscher, Viviana Ricardez, Carl J. Franklin, George J. Guillen

667–670
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Novel observations of arachnid predation on the Mountain Skink, Chalcides montanus Werner, 1931, and the Algerian Sand Gecko, Tropiocolotes algericus Loveridge, 1947, in Morocco

Mohamed Mousaid, Abdellah Bouazza

857-860
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An Eastern Ribbonsnake, Thamnophis saurita (Linnaeus, 1766), scavenging on a roadkilled Cuban Treefrog, Osteopilus septentrionalis (Duméril & Bibron, 1841), in Everglades National Park, Florida, USA

Sarah Payne, Eleanor Lane, Faith Dunlap, Madison Vasquez, Matthew Metcalf, Lisa M. McBride, Sarah Sherburne, Christina M. Romagosa, Amanda M. Kissel, Amy A. Yackel Adams, Mark R. Sandfoss

387–388
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Early maturation and high growth rates in an Eastern Treefrog, Hyla orientalis Bedriaga, 1890, population after the species’ expansion in Kaliningrad Oblast (East Baltic Sea region)

Artem A. Kidov, Roman A. Ivolga, Tatyana E. Kondratova

781-786
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Biofluorescence in the Madeiran Wall Lizard, Teira dugesii (Milne-Edwards, 1829)

José Câmara, José Jesus

679–682
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Expanding the known range: Spalerosophis diadema diadema (Schlegel, 1837) in the temperate western Himalayan region, India

Muzaffar A Kichloo, Sudesh Kumar, Neeraj Sharma

671-672
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Further herpetofaunal records from Kastos and Kalamos islands, with comments on the occurrence of Emys orbicularis (Linnaeus, 1758) and Podarcis ionicus (Lehrs, 1902) in the adjacent Meganissi, Ionian Islands, Greece

Elias Tzoras, Labros Logothetis, Panagiotis Drakopoulos

963-967
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Bioacoustics of the False Toad Pseudobufo subasper Tschudi, 1838 from Southeast Asia

Renaud Boistel, Olivier S.G. Pauwels, Amadeus Plewnia

913–919
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First report of juvenile fish (genus Geophagus) in the diet of the neotropical caecilian Potomotyphlus kaupii (Berthold, 1859)

Darlison Chagas-de-Souza, Tássio Alves-Coêlho, Elzamara C. Oliveira, Luana C. Cruz, Alfredo P. Santos-Jr.

845–847
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First recorded predation of a Mediterranean Centipede, Scolopendra cingulata Latreille, 1789, by a North African Ocellated Lizard, Timon pater (Lataste, 1880), in Chréa National Park, Algeria

Mustapha Aroudj, Youcef Hamidi , Mohamed Abbad , Yamina Mouas, Mostefa Benacherine, Cherifa Chaouia

985-988
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Bite, shriek, or intimidate: defensive behaviours of the Apatani Horned Toad, Xenophrys apatani Saikia et al., 2024, and the Bicoloured Frog, Clinotarsus curtipes (Jerdon, 1853), from India

Tage Tajo, A.N.D. Akalabya Sarmah, Upadhyaya K. Radhakrishna, Sonali Garg, Sathyabhama D. Biju

999-1004
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An unexpected urbanite: first record of a Plateau Toad, Anaxyrus compactilis (Wiegmann, 1833), from a developed site in Tlaxcala, Mexico

Maribel Méndez-Tepepa, Kevin I. Medina Bello, J. Erick Gómez-Campos

1005–1007
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A new observation of ophiophagy in the Madagascar Black-backed Smooth Snake, Liophidium vaillanti (Mocquard, 1901), in the Kirindy Forest, western Madagascar

Rina H. Andriamihantason, Peter M. Kappeler, Fanomezana M. Ratsoavina

1009–1010
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Passive acoustic monitoring protocol reveals new occurrence data for Eleutherodactylus bartonsmithi Schwartz, 1960 in eastern Cuba

Mariam Curbelo Cruz, Sergio L. del Castillo Domínguez, Leandro Galano Machado, Roberto Alonso Bosch

1021-1025
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Observations on nocturnal activities of the Dumeril’s Monitor, Varanus dumerilii Schlegel, 1839, in Borneo, Malaysia

Zichen Qiao, Yuchen Hu, Ingg Thong Law, Ing Sind Law

1027-1030
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Morphometrics aid identification of non-native iguanas to improve conservation of native Saba Green Iguanas, Iguana iguana (Linnaeus, 1758)

Matthijs P. van den Burg, Jeroen Kappelhof, Adolphe O. Debrot

1031-1037
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Range extension and first record of Dendropsophus studerae (Carvalho-e-Silva, Carvalho-e-Silva, and Izecksohn, 2003) in the state of Pernambuco, northeastern Brazil

Igor J. Roberto, Yuri Gomes, Rodrigo Leite, Pedro Barbosa, Daniel M. de Lima, Ednilza Maranhão dos Santos

1039-1042
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Comments on the retraction of an article by Faiz et al. (2017) on the putative occurrence of a King Cobra, Ophiophagus hannah (Cantor, 1836), in Pakistan

Hinrich Kaiser

1043–1046
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Feeding on different life stages of the Common Spadefoot Toad, Pelobates fuscus (Laurenti, 1768), by the Medicinal Leech, Hirudo medicinalis

David C. Broek, Mick Vos, Jöran Janse

1047–1049
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Beyond the asymptote: a near-complete reptile inventory in the Miombo woodland and on Mount Morué, Mocuba Municipality, with a major range extension for the lizard Nucras boulengeri Neumann, 1900

Avelino R. Miguel, Berta I. Sitole, Nando A.M. Calonga, Celso H. Duarte, Dolescêncio Armando, Célia J.A. Nanvonamuquitxo, Hinrich Kaiser

1051–1071
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Gut and faecal microbial diversity in a Japanese lotic-breeding salamander, Hynobius boulengeri (Thompson, 1912)

Nagisa Tasaki, Kanto Nishikawa, Kohei Nakamura, Hirotoshi Sato

1073–1077
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Death by geochemistry: sudden mortality among larvae of Korean Clawed Salamander, Onychodactylus koreanus Min et al., 2012, in Hwanseon Cave, South Korea

Daesik Park, Jaebeom Jeong, Min-Woo Park, Jongsun Kim, Hahyun Nam, Narae Joo, Jaejin Park

1089-1093
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Egg predation of a rhacophorid foam-nest frog (Chiromantis cf. rufescens) by a colubrid tree snake (Dipsadoboa sp.) in Salonga National Park, Democratic Republic of the Congo

Gabriel Badjedjea, Janis Czurda, Franck M. Masudi, Felix F. Kihambu, Vaclav Gvoždík

1095-1099
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Deimatic anti-predator behaviour in Uroplatus henkeli Böhme & Ibisch, 1990 during predation by Ithycyphus miniatus (Schlegel, 1837) on Nosy Be, Madagascar

Sam Hyde Roberts

1101-1103
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The diet spectrum of Pelophylax waterfrogs from mainland Greece, including novel data for the Epirus Waterfrog, Pelophylax epeiroticus (Schneider et al., 1984)

Petr Papežík, Michal Fečák, Michal Benovics, Adam Javorčík, Jozef Balcerčík, Peter Mikulíček

1105-1111
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Observations on predation by Mimophis mahfalensis (Grandidier, 1867) in Andranomena Special Reserve, Madagascar

Nirindrainiarivony P.H. Djadagna-Ahy, Achille P. Raselimanana

1113–1115
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